The 50,000 sq ft (4,645 sq m) supermarket will be the most environmentally friendly of its kind and is expected to reduce the store's carbon footprint by half, compared with a conventional supermarket of comparable size.
Features of the eco-store include a sustainable timber frame, micro wind turbines, solar cells, rainwater harvesting - collected from the roof and re-used in store - and energy-efficient refrigeration and bakery ovens.
There was also a significant increase in the use of recycled, recyclable or sustainable materials in the store's construction.
Plans for the eco-store were announced by Tesco chief executive Sir Terry Leahy in May, as part of the group's 10-point Community Plan. Leahy also revealed the launch of a£100 million fund for investment in environmental technology.
Tesco company secretary Lucy Neville-Rolfe said: 'The Wick store is a great step forward for Tesco. It brings together the latest construction, engineering and sustainable technologies.'
Carbon Trust director of delivery and external relations Dr Garry Felgate said: 'Tesco's new store at Wick is a great example of how businesses can incorporate innovative ideas into new buildings and create stores with a low carbon footprint.'
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